My husband, Ulrich Loening, who has died aged 94, was a lecturer first in the botany and then in the zoology departments (now the molecular plant sciences and biological sciences departments) of Edinburgh University. He engaged in fundamental research and made significant contributions to the developing science of molecular biology.
He had always had a great interest and concern for the natural world. After the establishment by Conrad Waddington, a professor of genetics, of the school of the man-made future at Edinburgh, Ulrich became actively involved in environmental and green issues.
